Beatrix Keener

Slithering out of the late-1980s Copper Age in the pages of Captain America, Beatrix Keener is a Marvel villain with a venomous pedigree — a card-carrying member of the Serpent Society, one of the most entertainingly sinister criminal collectives in the Marvel Universe. Debuted in 1988 under the sharp creative hands of Mark Gruenwald and Kieron Dwyer, she's kept dangerous company across the decades, sharing panels with fellow Society members like Coachwhip and Puff Adder as well as a colorful rogues' gallery of supporting players. With appearances stretching all the way to 2025 and two key collector issues to her name, Beatrix is a rewarding deep-cut discovery — the kind of persistent, underrated presence across Captain America, Spider-Woman, and The Amazing Spider-Man that reminds you just how richly populated the Marvel Universe really is.